Toulouse are embarking on their first ever season in Europe's premier competition and will host last season's finalists in the third qualifying round on Wednesday.

And Elmander's strike will see them in buoyant mood having quickly recovered from their opening day 3-1 defeat to Valenciennes a week ago.

"All week we worked on rediscovering our virtues, our desire, our solidarity, and that was the case this evening," said Toulouse coach Elie Baup, who was nevertheless worried that his team would struggle to recover from the match in time for Wednesday.

"It was very hot today," he added. "It won't be easy to play again in the same conditions on Wednesday."

Anyone hoping that this reverse may herald a changing of the guard in French football may be bitterly disappointed, though.

The six-in-a-row champions never gave anyone else a chance last season, almost wrapping up the title by Christmas, but this defeat owed much to the dismissal of Swedish midfielder Kim Kallstrom after just 34 minutes for stupidly spitting in the direction of the fourth official.

Until then Lyon had been the dominant force and there was nothing to suggest they will not canter to another French title again this season.

Lyon were left disappointed at coming so close to holding out for the point, but ultimately coming up short.

"It's tough when you play 10 against 11 for the whole match and you concede a goal at the end - it's really disappointing," said midfielder Mathieu Bodmer.

"Toulouse put a lot of physical pressure on us. We did what we could, we tried to earn a point but we didn't manage it."

Brazilian playmaker Juninho tested Toulouse goalkeeper Nicolas Douchez from a free-kick on 19 minutes and on 33 minutes Karim Benzema fired narrowly wide.

But the red card changed the complexion of the match.

Toulouse started to make the extra man count after the break as Andre-Pierre Gignac came close before defender Mauro Cetto thuded the bar with a header on 54 minutes.

Lyon remained dangerous as Abdul Kader Keita hit the post before Bryan Bergougnoux cleared the danger with his keeper beaten.

Lyon looked good for a point despite chances for Gignac and substitute Francois Sirieux until Elmander's exquisite finish with the outside of his boot in the last minute of regulation time.

Marseille remain winless and goalless despite the return of Samir Nasri after a second straight 0-0 draw, this time at home to Rennes.

Bordeaux - under new coach and former Manchester United, Barcelona, Marseille and Inter Milan centre-back Laurent Blanc - are the early high-flyers with six points from their two games having won 2-0 at Auxerre, although they had two Toifilou Maoulida penalty misses to thank for that win.

Brazilian midfielder Wendel gave Bordeaux the lead on the stroke of half-time and he added the second seven minutes after the break.

Nancy, who beat Caen 1-0, and Le Mans, who won 3-1 at Sochaux, also have a perfect six points from six. St Etienne and Lorient follow in fourth and fifth places after victories over Valenciennes (3-1) and Monaco (2-1) respectively.

Nice secured their first points of the season as a late Vincent Hognon goal sunk 10-man Strasbourg.

The big match of the weekend sees Guy Roux's Lens entertain Paul Le Guen's Paris St Germain on Sunday while Lille travel to Metz.